The shannon wiener diversity index is one measure that we will use to try to draw information from samples in the field. Diversity can be partitioned into two independent components. Lets compute the shannonweiner diversity index for the same hypothetical community in the previous example. Species diversity, community structure, and distribution. A measure derived from information theories that were developed by claude elwood shannon 19162001 and norbert wiener. Shannonwiener index is defined and given by the following function. Nov 27, 2015 an example of the shannonwiener diversity index for science students. Jan 06, 2011 demonstration of how to calculate the shannonwiener diversity index in microsoft excel. Mar 19, 2018 it apparently the shannonwiener index was not closely associated to reflectance, as the blackblue curves indicate a high shannonwiener index didnt merge together fig. Pdf shannonwiener diversity index has been used extensively in environmental studies to estimate the species.
For categorical data, there is no mean or median and so the measures of variation described in measures of variability cant be used. The shannonweiner species diversity index association for. Instead we use a measure of the distribution of observations among the categories. Dominance indices are weighted toward the abundance of the commonest species. Can i compare shannonwiener diversity indices between. Shannonwieners index of diversity was computed using the following formula. Biased richness and evenness relationships within shannon.
In fact, in the late 1940s shannon had built on the work of wiener. Engels department of zoology, state university of ghent, belgium in a low diversity brackish water habitat the diversity of the copepod community is best measured with the shannon wiener information function and its evenness by an. Natural resource biometrics measures of diversity shannon. There are a number of diversity indices which can be used to summarize the diversity in a community. Many indices for measuring species diversity have been proposed. Shannonwiener index is the most widely index which used in marine biodiversity. This comparison can be between two different habitats or a comparison of one habitat over time. Apr 11, 2003 on some occasions it is called the shannonwiener index and on other occasions it is called the shannonweaver index. The simpson index is mathematically simple for students to grasp.
Statistics shannon wiener diversity index tutorialspoint. The purpose of this analysis was to empirically model and graphically illustrate the numerical relationships between richness s, 435 species and evenness e with respect to shannonwiener index h. A diversity index is a quantitative measure that reflects how many different types such as. The shannonwiener diversity index h has been used to characterize grooming relationships among adult female primates.
Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. Application of shannons index to study diversity with. How to calculate shannon wiener diversity index definition. Perhaps the most celebrated index is the shannonwiener diversity index. On some occasions it is called the shannonwiener index and on other occasions it is called the shannonweaver index. They are just indices of diversity, in the same way that the diameter of a sphere is an index of its volume but is not itself the volume. For our uses, this order could be characterized by the number of species andor the number of individuals in each species, within our sample plot. Shannonwiener index, also known as shannon entropy, is a measure of information content and unpredictability in ecological studies margalef, 1958. The calculator uses the following formula to calculate the shannon wiener diversity index. Simpson diversity and the shannonwiener index as special. The shannon index has been a popular diversity index in the ecological literature, where it is also known as shannons diversity index, the shannonwiener index, the shannonweaver index and the shannon entropy. Woody species composition, structure and diversity of. Based on shannon wiener diversity index, the mannwhitney u test showed a significant difference between the two habitats u 2. Now from the alpha diversity tab choose the shannon wiener test we could.
Thus, when conducting simple statistical analyses of a specific effect of disturbance on diversity, the choice of index does not appear particularly important. Diversity indices provide more information about community composition than simply species richness i. Shannonweiner index assumes that individuals are randomly sampled from an independent large population and all the species are represented in the sample. Shannon weiner index assumes that individuals are randomly sampled from an independent large population and all the species are represented in the sample. In this article, a link is noted between a common family of diversity indices and non. The shannon index is a measurement used to compare diversity between habitat samples. Species diversity considers both the number of species richness in the community and their relative abundance evenness or equability. Bacterial diversity was positively correlated with plant diversity. Here, we will explore two measures of biodiversity. Pdf on jan 1, 2006, kathleen a nolan and others published beachcomber biology. Plant diversity richness index and the shannonwiener diversity index was significantly and positively correlated to a greater degree with moisture in the topsoil 05 cm layer than in the subsoil, and was also closely correlated to the humidity index.
The shannon wiener diversity index h has been used to characterize grooming relationships among adult female primates. Introduction, simpsons index and shannonweiner index. Lake superior, zooplankton, diversity index, regionalization. We will then consider one measure used to compare diversity among communities, jaccard. A diversity index is the measure of species diversity in a given community. The shannonweiner index is most sensitive to the number of species in a sample, so it is usually considered to be biased toward measuring species richness. This measure of equitability compares the observed shannonwiener index against. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. These metrics are a measure of the likelihood that a selected pixel will have the same land cover class as the adjacent pixel. Shannons diversity index real statistics using excel. C the more species you have, the more diverse the area. The shannonwiener diversity index is one measure that we will use to try to draw information from samples in the field.
Assessment of plant species diversity based on hyperspectral. This index is widely employed to measure biological diversity magurran, 2004. We suggest that at the very least, authors should define. B based on the species richness the number of species present and species abundance the number of individuals per species. Calculate the shannon diversity index and evenness for these sample values. According to local occurrence status, there were 12 species ranked as very abundant, 19 abundant, 62 very. We considered elevation as an independent variable for both amphibians and reptiles and calculated species richness or abundance as a dependent variable. The actual formula for the shannonweiner diversity index is. In particular, for a random sample, we can use shannons index of diversity aka as shannonweiners index, which is. Richness and evenness the heterogeneity of a landscape can be quantified using the shannon measures for richness and evenness.
Shannon index is a commonly used diversity index that takes into account. A tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. This tutorial explains how to calculate the shannon wiener diversity index and evenness. The shannon evenness index ranges from 0 when one species is dominant to 1 when all species are equally abundant. Introduction in recent years, there has been a rapidly increasing interest in the effects of species richness on community productivity.
Dpsi log pi, where s is the total number of species in the sample, i is the total. Brief outlines of the two types of diversity indices of biodiversity are discussed in this article. The measure was originally proposed by claude shannon to quantify the entropy uncertainty or information content in strings of text. Diversity indices like the shannon entropy shannon wiener index and the ginisimpson index are not themselves diversities. Monitoring location outside industrial area site2 revealed slight higher diversity with shannon and wiener species diversity index of 0. Effects of precipitation changes on soil bacterial. To make comparisons among different groups, the value of h has been divided by the maximum value it can reach for a particular group size. A tribute to claude shannon 19162001 and a plea for more. Pdf modification of shannonwiener diversity index towards. A a diversity index is a mathematical measure of species diversity in a given community. Pdf a comparative study of fish assemblage and diversity. The shannon diversity index h is another index that is commonly used to characterize species diversity in a community.
Shannonwiener index of diversity information index a measure used by ecologists when a system contains too many individuals for each to be identified and examined. Effects of precipitation changes on soil bacterial community. A diversity index is a mathematical measure of species diversity in a community. Shannonwiener index of diversity information index source. First, let us calculate the sum of the given values.
The underlying mathematical theory relates to statistics probabilities, multivariate analysis, cluster analysis etc. Shannon index the idea behind this index is that the diversity of a community is similar to the amount of information in a code or message. Plant diversity was significantly influenced by the differences in soil water storage in the top layer 05 cm but not in the deeper layers. It measures the rarity and commonness of species in a community. A measure used by ecologists when a system contains too many individuals for each to be identified and examined. Shannon to quantify the entropy uncertainty or information content in strings of. The shannon index has been a popular diversity index in the ecological. An example of the shannonwiener diversity index for science students. The maximum diversity hmax of a sample is found when all species are equally abundant. Key wordsdiversity indices, shannon index, shannon wiener index, shannon weaver index, species richness, species diversity. Demonstration of how to calculate the shannonwiener diversity index in microsoft excel.
The degree of uncertainty of predicting the species of a random sample is related to the diversity of a community. Measuring diversity has been of historical significance and due to the obvious declines in habitat diversity this data is still valuable. This makes the shannon index and the simpson diversity or gini coefficient special cases of a more general index. Shannon wiener diversity index is a term in biologyecology. In this example, the first sample would be considered more diverse. Find shannons index of diversity and index of relative diversity for a random sample of 25 observations distributed among five categories as shown in range b4. Sample values s 60,10,25,1,4 number of species n 5. Posted on 201021 20170904 author bpmsg categories online tools, other articles tags berger parker, calculation, excel, shannon entropy, shannon weaver index, shannonwiener index, simpson index diversity as business kpi alpha and beta diversity.
There have been numerous attempts to create compound indices that combine measures of richness and abundance. These values were then subjected to a model to develop statistically homogeneous zones, and to determine the temporal and spatial variability of the diversity values. Dpsi log pi, where s is the total number of species in the sample, i is the total number of individuals in. Aug 12, 2015 a tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. The shannon wiener diversity index was calculated for the total zooplankton community found in lake superior in 1973, as well as for its components ie. A widely used dominance index is simpsons diversity index. The concept of diversity is well introduced in ecology, economy and information theory. A tribute to claude shannon 19162001 and a plea for. Shannonwiener index unlike the simpson index, h is interpreted that the higher the score the more diverse.
Foremost among these are the shannons diversity h and simpsons diversity d 1 indices table table1, 1, which differ in their theoretical foundation and interpretation magurran 2004. Other researchers were not content with this state of affairs, and. We suggest an explanation for this dual use of terms and in so doing we offer a tribute to the late claude shannon who passed away on 24 february 2001. A greater number of species and a more even distribution both increase diversity as measured by h. If i obtain the shannonweiner diversity index as 2. This diversity measure is based on information theory. The use of the shannonwiener diversity index to delineate the. Key wordsdiversity indices, shannon index, shannon wiener index, shannonweaver index, species richness, species diversity. Here p i is the proportion of total number of species made up of the ith. Species evenness was calculated using the shannon evenness index. H has its foundations in information theory and represents.
The shannon index measures the order within a community written as h. Aug 28, 2014 in analyses using only one diversity index, similar significant effects of lui would have been detected using any of the indices we included, except e. A diversity index is a quantitative measure that reflects the number of different. Jan 26, 2012 a diversity index is the measure of species diversity in a given community.